1997: PhD MRC National Institute for Medical Research at Mill Hill, London (UK). "Cis-regulatory mechanisms of the Hox genes in mouse development". Lab of Dr. Robb Krumlauf. 1997-1998: Postdoc on Xenopus development, University of Chile. 1998: MRC Human Genetics Unit, Edinburgh. Postdoc on computational approaches to study mouse limb development. 2001: Development of a 3D optical imaging technique and introduction of the term "Optical Projection Tomography", commercialised under the name Bioptonics. 2003: Group Leader in Edinburgh. 2006: Senior Group Leader at the Centre de Regulació Genòmica, Barcelona (Spain). 2011: Acting Coordinator of the EMBL-CRG Systems Biology Program. 2014: Coordinator of the EMBL-CRG Systems Biology Program.