Education 1978-82: BSc Biochemistry, Stirling University 1982-85: PhD Biochemistry, Oxford University 1985-90: Post-doct. Fellow, Biochemistry Department, Stanford University Research & professional experience 7/1/90-7/1/95: Assistant Prof., UC San Diego, Biology Dep. 7/1/95-7/1/99: Associate Prof. (tenured), UC San Diego, Biology Dep. 7/1/99-10/31/2008: Prof., UC San Diego, Cell & Developmental Biology Dep. 7/1/98-7/1/2003: Adjunct Member, Mario Negri Sud Research Institute, Italy 1999-2000: ICREA Visiting Scientist, Univ. of Barcelona, Spain 2007-Present: Chair, Cell & Developmental Biology, CRG, Spain Awards and Honours Secretary Oxford University Biochemical Society (1982-84); Pirie-Reid Scholar, Oxford University (1982-85); Damon-Runyon Walter-Winchell Post-doct. Fellow, Stanford University (1985-87); American Cancer Society Senior Post-doct. Fellow (1988-90); Basil O'Connor Scholars Award (1992-95); EMBO Elected Member (2009); ASBMB Merck award (2013).